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​The New Guinea Highland Wild Dog
The Highland Wild Dog (HWD) is one of the rarest and most ancient canids currently living, potentially our best example of a proto-canid and is truly a living fossil.  It is the apex predator of New Guinea and what many think is one of the most important canids in existence.  The HWD may be the missing link species between the first early canids and the modern domestic dog.
The discovery and confirmation of the HWD for the first time in over half a century is not only exciting but an incredible opportunity for science.  The NGHWDF is so very proud to be able to bring you the following story and information.  The images that appear on this page are of actual HWDs photographed in their native New Guinea habitat during the 2016 expedition.  We hope you enjoy them as much as we enjoy bringing them to you.

Please join us for discussion of the images, information and expedition on  The Den Site Blog Page.

​​Meet White Cheek Girl, a pregnant female seen in several photos.

​​​new guinea highland wild dog foundation, inc

Expedition 2016

Re-Discovery

Despite anecdotal reports and even two intriguing but unconfirmed photographs in recent years, many feared the New Guinea Highland Wild Dog had become extinct through loss of habitat and contamination by Village Dogs. 

The 2016 Expedition was able to locate, observe, gather documentation and biological samples, and confirm through DNA testing that at least some specimens still exist and thrive in the highlands of New Guinea.

Discovery

In September 2016, Mac McIntyre was on an ecotour in Papua Province when he encountered the University of Papua (UNIPA) rapid survey team searching for Highland Wild Dogs (HWDs).  Invited to join them to view a den site and search for dogs in the remote areas adjacent to Puncak Jaya (Carstensz Pyramid), Mac removed his shoes to cross a waterway and ascend the slopes leading to the target site.  After viewing the den site and exploring at elevation but failing to find dogs, Mac and the UNIPA team began descending the muddy mountain trail. 

Mac’s mood was as grey and damp as the weather.  His trip was nearly over and while he had been able to investigate and collect samples from some Village Dogs in the traditional Village of Banti, he had yet to see any Highland Wild Dogs, despite having heard stories of them being present in the region.  Still barefoot, Mac glanced down to navigate the rain soaked ground and noticed something odd.  There in the mud, next to his bare footprint from the ascending trip, were the unmistakable fresh pawprints of a canine.  As Mac stared down in awe at the pawprints following his own up the trail, he knew they could only belong to an HWD.  The HWD had found ​him.  Realizing that this elusive but curious canid had followed him up the trail, Mac could barely contain his excitement and immediately called the find to the attention of the UNIPA team.  

​Moment of Discovery

New Guinea Highland Wild Dogs, called Anging Penyani in the Bahasa Indonesian language, which means "Dog That Sings", may be the same as or a progenitor of the rare, captive New Guinea Singing Dog populations and also a probable ancestor of the Australian Dingo.  And despite stories, anecdotal reports and two intriguing but unconfirmed photographs, they have not been  documented with certainty in their native range in over half a century. 

Given the remoteness of the region and the fact that there are no permanent human inhabitants or village dogs in the area, any evidence of dogs could most l ikely be attributed to the HWD.  Trail cameras were immediately deployed throughout the area to monitor bait sites set up to attract the dogs.  Over the next two days, the cameras captured over 140 images of HWDs living at altitude (4500 m) on Puncak Jaya, just adjacent to the immense Lorentz National Park.  The team would soon discover that because Grasberg Mine and PT Freeport Indonesia (PTFI) have taken special environmental stewardship measures to protect the austere, remote area and ecosystem around the mine, they had inadvertently created a sanctuary in which the HWD could thrive.

Lady Foot, the HWD that followed Mac upslope and left her tracks near his

The HWD represents the apex predator for New Guinea.  Despite this, scientists know virtually nothing about the HWD and their status in the wild, in fact, some believed the HWD to be extinct.  The dogs were first described by Charles Walter De Vis when he collected a specimen at about 2400 meters on Mount Scratchley, Papua New Guinea, in 1897.  In 1956, Albert Speer and J.P. Sinclair obtained a pair of dogs from the Lavani Valley in the Southern Highlands Province, Papua New Guinea.  The pair was sent to Sir Edward Hallstrom for inclusion in his animal research center in Nondugi, Papua New Guinea, then to the Taronga Zoo in Sydney, Australia, where they would be examined by Ellis Troughton in 1958, at which time they were classified as a distinct species, Canis hallstromi, in honor of Sir Hallstrom.  The Taronga Zoo specimens and five additional specimens obtained in 1976 from Papua Province would be the foundation stock of the captive variant of dog now commonly referred to as the NGSD with a population estimated at about 200 worldwide, though the actual number is unknown.  Due to largely unregulated breeding and lack of a registry combined with a lack of HWD specimens until now, it is unclear how closely the captive specimens adhere to the phenotype, genotype and ecology of the HWD.  The current taxalogical classification for the NGSD is in a state of uncertainty and the subject of some controversy.

In addition to deploying trail cameras, the team was also able to observe and document HWDs in the area first hand, with Mac now serving as scientific advisor.  Fecal samples were also collected for testing, and early findings from the DNA analysis performed by Dr. Benjamin Sacks at UC Davis showed the HWD sample possessed the A29 haplotype, which is consistent with dingo and NGSD.  These samples were compared to the Village Dog samples collected by McIntyre at Banti.  The village dogs samples contained the B52 haplotype, indicating that population is an outgroup with the most likely explanation being importation (see VILLAGE DOGS .)  DNA analysis remains ongoing.

The Highland Wild Dog

Prior to the 2016 expedition, there had been only three possible sightings of the HWD in its native range of New Guinea and there have been only two attempts to study the species in the wild. The first was when Robert Bino conducted a field study in 1993 and then James K. McIntyre in 1996.  Neither was able to observe any animals but found what they presumed to be HWD scat, pawprints, urine marks and predations, and McIntyre reported hearing vocalizations, but with Village Dogs presenting similar or identical spore and no way to verify a genetically pure specimen, the presence of HWDs could not be confirmed and was in question due to increasingly overlapping habitat with Village Dogs and a lack of any current anecdotal reporting.  In recent years there have been only three reputable sighting reports; One by Flannery in 1989, one by Kristofer Helgan working at Lake Tawa in the Kaijende Highlands in 2005 and the third when Tom Hewitt, Director of Adventure Alternatives Borneo, managed to photograph a dog resembling an HWD at a distance in the Star Mountains of Papua Province, near the base of Puncak Mandala, in 2012.   (See  THE HEWITT PHOTO  .)

​Photo evaluators named this pretty girl Black Back Mom

DNA analysis of NGSDs has established them, and hence the HWD, as the most ancient, primitive, proto-domestic canid currently living. The fossil record indicates the species established itself on the island at least 6,000 years ago, believed to have arrived with human migrants, however new evidence suggests they may have migrated independently of humans (Wilton, Cairns 2016 - see LIBRARY ). While the taxonomy and phylogenetic relationships with related breeds and Australian Dingoes is currently controversial and under review for both NGSD and HWD, the scientific and historical importance of the HWD remains critical to understanding canid evolution, canid and human co-evolution and migrations, and human ecology and settlement derived from the study of canids and canid evolution..   Determination of status of wild specimens to examine populations, habitat, range, prey species and predator/prey relationships, ecosystem position and impact, ecology, and human interaction or threat is of critical importance, along with species conservation.  In short, the HWD is a “missing link” and foundational species with a key primary position in the evolutionary trajectory of all canids, especially those of Asian origins; it represents canids in a pristine, prototype state before and as canids began being domesticated. Further exploration is sorely needed and the NGHWDF is in the planning stages for a 2017 expedition.

​Black Back Mom's three plump pups, Two Socks, Lil Red, and Markie.  The NGHWDF Field Research team hopes to locate and identify these pups during the upcoming 2017 Expedition.

​Expedition
The 2016 expedition was conducted between 3460 and 4400 meters a.s.l. along the Sudirman Range near Pucak Jaya in Papua Province, Indonesian New Guinea.  The remote areas adjacent to the Grasberg Gold Mine (4°3’10” S, 137°6’57” E) consisted of alpine meadows (3900 – 4170 meters a.s.l.) with a shrub layer over mixed close ground cover consisting of grasses and herbs, and rocky barrens, which were the highest regions just below the glaciers, consisting of barren rocks largely devoid of vegetation apart from lichens and mosses.  Only a few bird and mammal species have adapted to these harsh circumstances. High mountain lakes and steep valleys dominate the landscape.  At the time of the expedition, the temperature fluctuated from 2 degrees Celsius at night to 11 degrees Celsius during the day (35 to 52 degrees F).  The mornings were generally clear but quickly deteriorated to lower temperatures with fog, mist, and light to heavy precipitation for the remainder of the daylight hours and often through the night. 
Attractants, lures and baits included:

  • Fresh estrus urine from captive NGSDs (collected August – September 2016)
  • Fresh cus cus urine
  • Evanesce Predator Curiosity Lure (beaver castor, mink gland secretion, skunk essence)
  • Karak Coyote Gland Lure (red coyote glands) and coyote gland scent obtained from hunters and hunting supply stores
  • Sandhill Magik Predator Paste bait (20 years plus aged bobcat meat)
  • Cooked chicken, fish and chicken bones obtained from the mine cafeteria

The team also used electronic game calls at all locations to broadcast the vocalizations of male coyotes, female coyotes, coyote howls, and coyote pup distress cries (canis latrans).  Despite belonging to a different species, the team hoped the coyote vocals would elicit a curiosity or territorial response, stimulate return vocalizations, or encourage HWDs to investigate, bringing them into proximity to the team for observation.  Coyote calls were successful in eliciting return canid vocalizations only twice, from the hills adjacent to the downslope town of Tembagapura.

​​​Images of Den Site

The research team also conducted scouting excursions to examine how the HWD was using the area, which included examination/documentation of an active den site located 2 km above and behind the mine. They were also able to observe HWDs along the roadways near the mine. The team collected twenty three fecal samples (scat) varying in age and degree of degradation from remote locations between 3460 and 4150 meters a.s.l.  Due to the remoteness of the location and the fact that the HWD is the largest mammal, as well as the apex predator inhabiting this region, it could be safely concluded that samples collected could be attributed to the HWD (which was later confirmed via DNA testing).  Some samples were examined visually and manually in the field while others were preserved for DNA analysis.

​Observations and Early  Findings

The 2016 expedition confirms that the HWD is present in its native and historical range, and has not been extirpated or become extinct as was once feared.  Dogs were noted as solitary individuals and in social groups of two, three and four individuals.  Individuals and groups consisted of adults of both sexes, pregnant and lactating females, and mothers with pups. Dogs were active during both daylight and nighttime hours when the weather was clear and without precipitation.

 Based on the photographic evidence obtained, it appears the dogs at these locations live in small social groups occupying distinct but possibly overlapping territories and travel routes.  Gender distribution within social groups could not be easily ascertained, nor if males are paired with individual females, groups of females or if they are independent, however their consistent presence in proximity to females that were pregnant or had already given birth may suggest they are part of a social group in some capacity.  It was also not possible to clearly define social structure and whether or not these groups are part of a larger social group.  Additional research is needed.
The expedition did not attempt to capture any dogs, so no data was collected for size or weight.  Analysis of the images and firsthand observations indicated that all the specimens appeared to fall within the normal physiological range of captive NGSDs.  NGSDs also exhibit some degree of sexual dimorphism, which seemed apparent in the HWD population, as well.  In general, the majority of the HWD population observed in the vicinity of the Puncak Jaya Range and Grasberg mine area exhibited phenotypical traits and morphology consistent with that of captive bred NGSDs in regard to proportions and shoulder height, length and estimated weight.  HWD specimens observed in the more remote regions appeared to be on the larger end of the spectrum and overall slightly larger than captive NGSD specimens, though hard data from captured HWD specimens is needed.

The HWDs observed presented as healthy and fit, possessing uniformly well furred coats, including dogs of obviously advanced age.  One female specimen, a mother with three pups (Black Back Mom - see above), was lean with sparser coat and visible ribs, but that’s consistent with wild canids engaged in the care and feeding of offspring that requires they devote significant energy and biological resources to their young. 
Coat coloration included cream, ginger, roan, black with white markings on chest, black with white markings in the Irish pattern, and darker roan or black with tricolor (black, white, tan) patterning.  White coloration presented in varying degrees in the form of blazes or sections on face, muzzles, shoulder and neck collars, chest, belly, forelegs and tips of tails.  Some of the cream, ginger or roan dogs exhibited overlays of black in the longer guard hairs, often over the back and/or on the dorsal side of tail.  Pups and young animals exhibited melanistic masking to varying degrees, such as is associated with being present at birth (“birth mask”) then phasing to white or prevailing pelage coloration.  Nose coloration was black with exception to one dog that displayed liver nose pigmentation. Feet were robust, compact, rounded to oval featuring long, well developed toes with pads that ranged from black to liver in color and toenails ranging from very light to black.

​A ginger pup and ginger pup with "Irish Marks", left; a black and tan male, center; and a roan pup with black guard hair overlay, right.

The tail is carried high over the dorsal posterior in a fish hook shape.  Some dogs had tails with bushy undersides while others had shorter, more uniform furring with a precaudal gland evident on many dogs.  In all HWDs observed, the underside of the tail was lighter in color than the rest of the coat, typically white or cream.  With exception to cream colored animals where it was difficult to tell, white tipping on the tail was present in varying degrees from sparse at the very tip to very prevalent extending 3 cm or more towards the base of the tail.  In all dogs observed, the ears were moderate in size, set on top of the head, erect and triangular in shape, with moderate interior furring.

​Tail carriage and ear set

In adult male HWD specimens where clear observation was possible, experts who examined the photos noted that the scrotum presented as moderately furred, compact and tucked tight to the body, in contrast to captive NGSDs in which case the scrotum is sparsely furred and the skin of the scrotum is loose, with the testicles and scrotum suspended pendulously and able to move freely beneath the body.   Intact male NGSDs are readily identified by their very visible scrotum; in the HWD images obtained, this feature was not observable. This could suggest that, similar to wolves, male HWDs are only fertile for a limited time during a breeding season, at which time testosterone increase causes the scrotum and testicles to “drop”, hanging freely below the body where descended testes would allow for cooler temperature necessary for sperm viability and act to as a virility signal for breeding.  Another explanation would be that in male HWDs, the scrotum is adapted to be compact and tight to the body to prevent injury in an austere environment or as a method to keep testes warm at cooler elevations when not actively engaged in seasonal breeding.  Further research is needed to confirm this trait is in fact present in all adult HWD males as well as examine exactly how, and more importantly why, it differs morphologically from captive NGSDs.

​Compact, furred scrotum in adult male HWD, left, compared to NGSD, right

Regarding the breeding season and reproductive behavior of the HWD, the research team identified pups ranging in estimated ages from to three to five months.  Two pregnant females very close to their whelping date, as determined by teat and vulva presentation as well as very heavy appearance and bulging sides, were also identified.  This would suggest a spring breeding season (also the dry season) from approximately May through September, considering location in the southern hemisphere.

​Other Noteworthy Observations

HWDs were also observed alongside roadways associated with Grasberg mine and near the vicinity of the mine.  The roadways are lined with piles of rock tailings that serve as barriers to prevent vehicles from driving off cliffs.   Dogs were observed and photographed standing or lying atop these rock piles and would exhibit normal, relaxed behaviors until a vehicle would begin to slow down or come to a stop in proximity to them, at which point the dogs would stand and become extremely wary and observant.  Photographic evidence had to be obtained quickly as prolonged presence of humans/vehicles or any attempt to exit the vehicles would cause the dogs to rapidly retreat down the back of the rock piles to the rock barrens or forest then disappear from sight. 

These same dogs appeared at irregular intervals and appeared to be habituated in the regard that they exhibited classic conservation of energy in scavenging and foraging for scraps amongst discarded box lunches provided for the mine workers.  As is generally accepted, habituated animals of any species are no less “wild” simply due to their propensity to scavenge human refuse, such as when wolves or bears frequent garbage dumps near human habitations.  This scavenging conservation of energy behavior exhibited by the HWDs instructed the research team to leave chicken bones and cooked fish in areas around the mine where dogs had been reported.  When the dogs eventually approached, the team observed that they would quickly snatch preferred foods, gulp down large amounts without chewing, then retreat rapidly to the security of the secluded, remote rock barrens, leaving containers and unwanted food, such as shrimp and rice, behind.

​An HWD watches warily, then makes a hasty retreat

The expedition team was directed to a den site where pups were born the year prior in May/June 2015.  The den was located in a remote area 400 meters behind and above the Grasberg Mine reclamation nursery station, where saplings are grown in pots until they can be planted.  The opening to the den was located in a rock formation at the base of a 3 meter high boulder providing access to an excavated tunnel approximately ¾ meter in diameter and extending downwards into the earth approximately 3 meters.  At the mouth of the den the research team observed the partial remnants of two fibrous straw pots (approximately½meter tall with a diameter width of ¼ meter) used at the mine nursery station to start seedlings.  The pots, which had been broken into pieces and shredded, had apparently been transported from the nursery station possibly 1 (one) km away to the den site by the pregnant mother as material to prepare and line her whelping den. 

​Pregnant White Cheek Girl urinates off trail while watched by her often seen companion, Fluffy Tail Girl.

As discussed previously, scats that could reasonably be attributed to the HWD were collected from various locations.  In instances where multiple scats were observed, they were typically within one 1 (one) meter of one another, suggesting the possibility that individuals or social groups use these sites as communal toileting areas.   In all cases, scats were deposited atop objects and manual/visual field examination of scat yielded bones of small mammals, such as mice and rats, as well as larger mammals, such as cus cus.  Bird and mammal claws, teeth, feathers, and hair, along with grass and gravel, were also present.  In instances where trails were obvious, scats were never deposited directly on the trail but rather were deposited just adjacent to the trail instead.

In addition to food scavenged from human refuse resulting from mine activity, the following mammals were identified as inhabiting the study area and suggest possible prey items of the HWD.

  • Tree Kangaroo
    • Dingiso, Dendrolagus mbaiso
  • Cuscus
    • Silky, Phalanger sericeus
    • Ground, Phalanger gymnotis
    • Mountain, Phalanger carmelitae
  • Ringtail Possums
    • Coppery, Pseudochirops cupreus
    • Pygmy, Pseudochirops mayeri
  • Rats
    • Glacier Rat, Stenomys richardsoni
    • Arianus’s Rat, Stenomys omlichodes
    • Moss Forest Rat, Stenomys niobe
    • Black Rat, rattus rattus

​Tree Kangaroo

​Pygmy Ringtail Possum, left, and Cuscus, right

​Coppery Ringtail Possum

​Black Rat

​Glacier Rat

​The Future

The author and supporters of this and future expeditions, along with numerous canid researchers and experts, consider the confirmation of the HWD, alive and thriving in its home range and ecosystem, to be one of the most important scientific breakthrough discoveries in recent years. 

In the past, research expeditions were hindered by the sheer vastness and austerity of the island along with political change.  With the 2016 expedition having successfully confirmed HWDs in the area around the periphery and slopes of Puncak Jaya (Carstensz Pyramid), to include the Grasberg Gold Mine vicinity, ongoing research at the mine is warranted and sustainable given the existing infrastructure support made available by PT Freeport Indonesia (PTFI), an affiliate of US based Grasberg Mine owner, Freeport-McMoRan Corporation.  PTFI’s commitment to sustain and protect the adjacent ecosystem through their long term, ISO 17025 Compliant Environmental Monitoring Program has created an ideal research environment in which to attract, hold and study the HWD while providing a stable sustenance resource platform for researchers.  It appears that the isolated, remote, high altitude location in conjunction with the extensive environmental safeguards and security measures set in place by PTFI in the vicinity of the mine and surrounding ecosystem have inadvertently created a sanctuary in which the HWD can thrive.  PTFI was deeply gratified to learn that their dedication to proactive environmental stewardship had created a safe haven for New Guinea’s only wild dog and apex predator, along with those who would study it, and has generously committed logistical support and funding to facilitate ongoing or even full time research long into the future. Environmental representatives from PTFI will continue to work with researchers to develop additional infrastructure and management plans to sustain and improve this invaluable natural resource. 
It is important to understand that while Grasberg Mine represents human activity, it is still an isolated, unrefined, remote location.  The nearest human settlement is Tembagapura, a town dedicated as infrastructure for mine personnel, located 16.1 km down slope from the mine, followed by the indigenous Village of Banti. Workers do not reside in the mine area, rather they travel to the site on a daily basis to perform their duties, and they are prohibited from taking dogs into the area.  The terrain between the village and town and the mine site is arduous; it is highly improbable that village dogs from these locations would abandon readily available and abundant prime resources (food/water, shelter and mates) to make the difficult journey to the challenging ecosystems at higher elevations, only to compete with established HWDs for limited resources once there.  Further, the HWD specimens inhabiting the highlands of Puncak Jaya and the Grasberg Mine vicinity do not display any phenotypical traits or morphology suggesting any recent or historical village dog admixture, nor have there been any reports of village dogs being present in the region.
While it is true that additional sites throughout the whole of New Guinea where HWD sightings have been occasionally reported by remote villagers  should be explored to locate and study HWDs in locations other than Puncak Jaya/Grasberg Mine, given the readily available subjects and infrastructure in place, future expeditions to the site are planned.  Research and goals of future endeavors include the following.

  • Capture of live specimens to obtain biological samples and physiological data; attach satellite tracking collars and means of identification; secure live specimens to be removed to captivity for further ongoing study
    • Understand range, distribution, population size/density, social construct, dispersal factors and patterns
  • A LIDAR survey to identify additional sites and gather data on existing sites
  • Deploy camera traps, DNA traps and remote listening stations (to study vocalizations); establish local monitoring/maintenance support for ongoing data collection
  • Conduct study to determine lure, bait and scent attractant preferences
  • Dispense/disperse wafer bait vaccines to inoculate HWDs against canine pathogens, particularly rabies;  vaccinate Village and domestic dogs to promote goodwill and as a further measure to prevent disease transmission
  • Comprehensive genetics analysis and study to determine origin and evolutionary trajectory; examine relationship to AUDs and NGSDs; comparative study to examine migration routes and co-evolution with humans
  • Establish a dedicated onsite research team or station
  • Study reproductive anatomy, seasons and behavior
  • Study longevity, mortality factors and rates, litter size and neonate mortality rates and causes
  • Develop a comprehensive conservation and management plan
  • Study predator-prey relationships and the environmental impact/importance of the HWD (or absence of) as the apex predator on the whole of New Guinea
  • Examine ecosystem, eco niche, and human impact factors in regard to the species
  • Study the ethology/ecology of the species, develop a valid ecogram

​White Cheek Girl investigates a scent bait

​A pup pauses in front of the IR camera

This page contains only a tiny fragment of the research and subsequent knowledge to be gained for many years to come.  Future research will take place under the newly founded New Guinea Highland Wild Dog Foundation (NGHWDF).  New Guinea represents a pristine environment and ecosystem in which to study flora and fauna in its most primitive, unchanged state.  The HWD is the apex predator for New Guinea; it is imperative to understand this species in order to gage health and fitness of the entire ecosystem.  Further, the HWD phylogentic lineage is critical to the conservation of the Australian dingo and fully understanding the evolution and migration of canids and humans moving out of Asia.

​​HWD Image Gallery

​Use the arrows to scroll through the images, or click on the gallery to view in expanded mode with descriptions.

  1. The New Guinea Highland Wild Dog Gallery
    The New Guinea Highland Wild Dog Gallery
  2. White Cheek Girl, soon to deliver pups
    White Cheek Girl, soon to deliver pups
  3. White Cheek Girl and Fluffy Tail pause trailside to urine mark
    White Cheek Girl and Fluffy Tail pause trailside to urine mark
  4. Meet Skinny Girl, a very thin female
    Meet Skinny Girl, a very thin female
  5. Skinny Girl - Note the anomoly on her lower jaw
    Skinny Girl - Note the anomoly on her lower jaw
  6. We believe this is also Skinny Girl.
    We believe this is also Skinny Girl.
  7. Black Back Mom visiting a scent bait site
    Black Back Mom visiting a scent bait site
  8. Black Back Mom's Pups, Two Socks, Lil Red and Markie
    Black Back Mom's Pups, Two Socks, Lil Red and Markie
  9. BT Boy 1, A black and tan male
    BT Boy 1, A black and tan male
  10. An HWD on the roadside rock tailings
    An HWD on the roadside rock tailings
  11. The HWDs observed first hand were shy and wary
    The HWDs observed first hand were shy and wary
  12. BT Boy 2 approaches a scent bait
    BT Boy 2 approaches a scent bait
  13. BT Boy 2 with conspecfics at scent bait site
    BT Boy 2 with conspecfics at scent bait site
  14. A male pup explores the scent bait
    A male pup explores the scent bait
  15. A second pregnant female
    A second pregnant female
  16. Lady Foot, the HWD that found Mac
    Lady Foot, the HWD that found Mac
  17. Flick, A healthy, energetic roan pup
    Flick, A healthy, energetic roan pup
  18. An HWD performs scent acquisition
    An HWD performs scent acquisition
  19. Pregnant female joined by BT Boy 3
    Pregnant female joined by BT Boy 3
  20. Black and Tan Male
    Black and Tan Male
  21. Black Back Mom's pups, Lil Red and Markie, romp and play
    Black Back Mom's pups, Lil Red and Markie, romp and play
  22. An HWD den
    An HWD den
  23. An HWD den
    An HWD den
  24. Puncak Jaya Region
    Puncak Jaya Region
  25. Puncak Jaya Cloud Forest
    Puncak Jaya Cloud Forest
  26. Puncak Jaya Region
    Puncak Jaya Region
  27. PUncak Jaya Region
    PUncak Jaya Region
  28. The HWD Ecosystem in the Puncak Jaya Range
    The HWD Ecosystem in the Puncak Jaya Range
  29. The Cloud Forest Ecosystem of Puncak Jaya
    The Cloud Forest Ecosystem of Puncak Jaya
  30. Puncak Jaya Region, Papua Province, New Guinea
    Puncak Jaya Region, Papua Province, New Guinea
  31. The HWD Habitat of Puncak Jaya
    The HWD Habitat of Puncak Jaya
  32. The Puncak Jaya Region
    The Puncak Jaya Region
  33. The HWD Ecosystem of Puncak Jaya
    The HWD Ecosystem of Puncak Jaya